If Legacy uses Next Evolution to become immune to Melee damage, and a card played during the Villain turn destroys Next Evolution, is he still immune to Melee damage until the start of his next turn?
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He is indeed. Anytime a card states "Until the start of your next turn" or the like, it will remain in effect until the start of your next turn whether it be there or not. Now if cards like Inspiring Presence would be destroyed then the effect would end.